Table 2 List of input and output variables

T _i,j (t)

I

Mean of minimum temperature from October (year t − 1) to March (year t) (°C)

P _i,j

I

Pine density (km⁻²)

H _i,j

I

Human population density (km⁻²)

N _i,j (t)

O

Population density (km⁻²) during the winter [t − 1, t], but before winter mortality

E _i,j (t)

O

Population density after winter mortality, at the time of emergence (year t)

A _i,j (t)

O

Population density after adult dispersal (year t)