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Table 1 Overview of the description of phenology in terrestrial ecosystem models (TEMs)

From: Temperate and boreal forest tree phenology: from organ-scale processes to terrestrial ecosystem models

Model class

Model

Vegetation type

Application scale

Leaf onset

Leaf loss

Wood growth

Fine root growth

Reserve storage

Fine root mortality and turnover

Reference

NPP allocation

CASA-CNP

PFT

globe

satellite

satellite

start with leaf onset, ends with leaf fall (deciduous); year-round (evergreens). Coefficients modulated according to leaf phenology

start with leaf onset, ends with leaf fall (deciduous); year-round (evergreens). Coefficients modulated according to leaf phenology

no reservepool

proportionnal to FR mass

Wang et al. (2010)

NPP allocation

ED2.1

PFT

globe

satellite

satellite

start with leaf onset, ends with leaf fall (deciduous); year-round (evergreens). Coefficients modulated according to leaf phenology

start with leaf onset, ends with leaf fall (deciduous); year-round (evergreens). Coefficients modulated according to leaf phenology

out of the leaf growth phase

proportionnal to FR mass, modulated by temperature

Medvigy et al. (2009)

NPP allocation

ISAM

PFT

globe

Virtual leaf C balance

Temperature and Photoperiod threshold

allocation to wood and roots start once maximum LAI is reached

allocation to wood and roots start once maximum LAI is reached

no reserve pool

proportionnal to FR mass

Arora & Boer (2005)

NPP allocation

ANAFORE

Tree species (Fagus sylvatica, Quercus petraea, Pinus sylvestris)

stand

GDD and chilling

prescribed

onset defined relative to that of leaves, ends with leaf senescence onset

simultaneous to leaf growth

modulated according toleaf and woodphenology

proportionnal to FR mass

Deckmyn et al. (2006, 2008)

NPP allocation

CASTANEA

Tree species (Fagus sylvatica, Quercus petraea, Pinus sylvestris, Picea abies, Quercus ilex)

stand

GDD

CDD

cambium onset at budburst (diffuse-porous deciduous), according to thermal time (ring-porous deciduous and conifers). Cessation at a prescribed date

modulated according to leaf and wood phenology

modulated according toleaf and woodphenology

proportionnal to FR mass

Dufrêne et al. (2005),Guillemot et al. (2015)

Labile pool allocation

OCN

PFT

globe

GDD and chilling

Temperature threshold

Depend on the availability of reserve C and N, modulated by daily temperature and leaf phenology

Depend on the availability of reserve C and N, modulated by daily temperature and leaf phenology

Reserve (labile) pools are the primary pool for storagefollowingassimilation

FR shedding simultaneous to leaf shedding

Krinner et al. (2005),Zaehle et al. (2010)

Labile pool allocation

CASSIA

Tree species (Pinus sylvestris)

stand

GDD

prescribed

cambium onset and cessation according to thermal time

forced

Reserve (labile) pools are the primary pool for storage following assimilation

no mortality

Schiestl-Aalto et al. (2015)

Labile pool allocation

CLASS

Tree species (Populus tremuloides)

stand

GDD

GDD

Depend on the availability of reserve C and N, modulated by daily temperature

Depend on the availability of reserve C and N, modulated by daily temperature

Reserve (labile) pools are the primary pool for storage following assimilation

Shedding of all organs (including FR) depends on thermal time. All organs have the same thermal time requirement for shedding

Wang et al. (2001)

  1. A representative panel of models is presented, including TEMs with both “NPP” and “Labile pool” allocation schemes (see main text and Fig. 4). Only models that simulate allocation or growth at a daily time step are considered
  2. NPP net primary production, PFT plant functional types, FR fine roots, GDD growing degree-day, CDD cold degree-day